WoRMS source details
Read, Geoffrey B. (2011). A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900. Zootaxa. 2934: 39-52.
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Read, Geoffrey B.
2011
A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900
Zootaxa
2934: 39-52
Publication
Polychaeta dbase. Not open access. Full text link: http://www.mapress.com/zootaxa/2011/2/zt02934p052.pdf
A maldanid discovered on sand beaches of Banks Peninsula, South Island, New Zealand is newly described. Clymenura snaiko sp. nov. is of large size at over 250 mm length, with 18 chaetigers, two achaetous preanal segments with tori, unmodified dentate uncini in anterior chaetigers, a ventral glandular shield on chaetiger eight, a high cephalic rim notched laterally, and a deep anal funnel rimmed with 16 short subequal cirri and one longer ventral cirrus. The confused taxonomy of ventral-shield-bearing maldanids is reassessed and clarified. Leiochone Grube 1868 and Clymenura Verrill, 1900 are here treated as the valid members of Arwidsson’s (1906) tribe Leiochonini in Euclymeninae, and given emended diagnoses, while Imajima and Shiraki’s (1982) subfamily Clymenurinae and Clymenura subgenus Cly. (Cephalata) are considered invalid. Differentiating character states of the valid Leiochonini species are tabulated. Leiochone is restricted to six species, including the new combinations L. annulata (Mohammad, 1980) and L. japonica (Imajima & Shiraki, 1982). Clymenura is restricted to nine valid taxa, including Cly. snaiko sp. nov. and the new combination Cly. polaris lena (Averincev, 1990). These updated groupings exclude a further eight nominal taxa which mostly may be indeterminable. Leiochone species occur in inshore temperate to tropical Eastern Hemisphere waters, while Clymenura mostly occur in high North European latitudes and the northern Pacific region, usually offshore in deep water. The new species is the first record of a Clymenura from Southern Hemisphere waters, and the second member of the genus to be found in the intertidal.
New Zealand
Clymene ebiensis Milne Edwards, 1843 (status source)
Clymene urceolatus Leidy, 1855 (status source)
Clymenura Verrill, 1900 (status source)
Clymenura (Cephalata) Imajima & Shiraki, 1982 accepted as Clymenura Verrill, 1900 (status source)
Clymenura (Clymenura) accepted as Clymenura Verrill, 1900 (status source)
Clymenura (Clymenura) japonica Imajima & Shiraki, 1982 accepted as Leiochone japonica (Imajima & Shiraki, 1982) (new combination reference)
Clymenura aciculata Imajima & Shiraki, 1982 (status source)
Clymenura annulata Mohammad, 1980 accepted as Leiochone annulata (Mohammad, 1980) (source of synonymy)
Clymenura borealis (Arwidsson, 1906) (status source)
Clymenura clypeata (Saint-Joseph, 1894) accepted as Leiochone leiopygos (Grube, 1860) (status source)
Clymenura columbiana (Berkeley, 1929) (status source)
Clymenura gracilis Hartman, 1969 (status source)
Clymenura johnstoni (McIntosh, 1915) accepted as Leiochone johnstoni McIntosh, 1915 (status source)
Clymenura leiopygos (Grube, 1860) accepted as Leiochone leiopygos (Grube, 1860) (status source)
Clymenura longicaudata Imajima & Shiraki, 1982 (status source)
Clymenura polaris acirrata (Zachs, 1923) accepted as Leiochone polaris acirrata Zachs, 1923 (status source)
Clymenura polaris lena (Averincev, 1990) (status source)
Clymenura snaiko Read, 2011 (original description)
Clymenura tenuis (Day, 1957) accepted as Leiochone tenuis Day, 1957 (status source)
Clymenurinae Imajima & Shiraki, 1982 accepted as Leiochonini Arwidsson, 1906 (status source)
Leiocephalus parvus Quatrefages, 1866 (status source)
Leiochone annulata (Mohammad, 1980) (new combination reference)
Leiochone borealis Arwidsson, 1906 accepted as Clymenura borealis (Arwidsson, 1906) (status source)
Leiochone clypeata Saint-Joseph, 1894 accepted as Leiochone leiopygos (Grube, 1860) (status source)
Leiochone japonica (Imajima & Shiraki, 1982) (new combination reference)
Leiochone polaris (Théel, 1879) accepted as Clymenura polaris (Théel, 1879) (new combination reference)
Leiochone polaris lena Averincev, 1990 accepted as Clymenura polaris lena (Averincev, 1990) (new combination reference)
Leiochone singularis Gravier, 1906 (status source)
Leiochone tropica Mesnil & Fauvel, 1939 (status source)
Leiochonini Arwidsson, 1906 (status source)
Praxilla simplex Claparède, 1869 (status source)
Clymene urceolatus Leidy, 1855 (status source)
Clymenura Verrill, 1900 (status source)
Clymenura (Cephalata) Imajima & Shiraki, 1982 accepted as Clymenura Verrill, 1900 (status source)
Clymenura (Clymenura) accepted as Clymenura Verrill, 1900 (status source)
Clymenura (Clymenura) japonica Imajima & Shiraki, 1982 accepted as Leiochone japonica (Imajima & Shiraki, 1982) (new combination reference)
Clymenura aciculata Imajima & Shiraki, 1982 (status source)
Clymenura annulata Mohammad, 1980 accepted as Leiochone annulata (Mohammad, 1980) (source of synonymy)
Clymenura borealis (Arwidsson, 1906) (status source)
Clymenura clypeata (Saint-Joseph, 1894) accepted as Leiochone leiopygos (Grube, 1860) (status source)
Clymenura columbiana (Berkeley, 1929) (status source)
Clymenura gracilis Hartman, 1969 (status source)
Clymenura johnstoni (McIntosh, 1915) accepted as Leiochone johnstoni McIntosh, 1915 (status source)
Clymenura leiopygos (Grube, 1860) accepted as Leiochone leiopygos (Grube, 1860) (status source)
Clymenura longicaudata Imajima & Shiraki, 1982 (status source)
Clymenura polaris acirrata (Zachs, 1923) accepted as Leiochone polaris acirrata Zachs, 1923 (status source)
Clymenura polaris lena (Averincev, 1990) (status source)
Clymenura snaiko Read, 2011 (original description)
Clymenura tenuis (Day, 1957) accepted as Leiochone tenuis Day, 1957 (status source)
Clymenurinae Imajima & Shiraki, 1982 accepted as Leiochonini Arwidsson, 1906 (status source)
Leiocephalus parvus Quatrefages, 1866 (status source)
Leiochone annulata (Mohammad, 1980) (new combination reference)
Leiochone borealis Arwidsson, 1906 accepted as Clymenura borealis (Arwidsson, 1906) (status source)
Leiochone clypeata Saint-Joseph, 1894 accepted as Leiochone leiopygos (Grube, 1860) (status source)
Leiochone japonica (Imajima & Shiraki, 1982) (new combination reference)
Leiochone polaris (Théel, 1879) accepted as Clymenura polaris (Théel, 1879) (new combination reference)
Leiochone polaris lena Averincev, 1990 accepted as Clymenura polaris lena (Averincev, 1990) (new combination reference)
Leiochone singularis Gravier, 1906 (status source)
Leiochone tropica Mesnil & Fauvel, 1939 (status source)
Leiochonini Arwidsson, 1906 (status source)
Praxilla simplex Claparède, 1869 (status source)
Okains Bay for Clymenura snaiko Read, 2011
Holotype NIWA 60439, geounit New Zealand Exclusive Economic Zone, identified as Clymenura snaiko Read, 2011
Etymology
‘snaiko’ is coined to form a sinuous-sounding recombination of the letters of the type locality place name, and ... [details]
New combination
Suggested new combination by Jiménez-Cueto and Salazar-Vallejo (1998) on basis of pygidial morphology, but the ... [details]
Taxonomy
Leiochone was erected by Grube (1868), without formality, for Clymene urceolata, Leidy, and Cl. leiopygos Gr. In ... [details]
Taxonomy
Read (2011) maintained as Leiochone borealis distinct and confirmed recombination in Clymenura. Distinct from Cly. ... [details]
Type locality
Okains Bay, Banks Peninsula, South Island, New Zealand, 43.6946°S, 173.0641°E, 0 m [details]
