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Hendricks, M. G. J.; Gibbons, M. J. (2010). Two new nematode species from Saldanha Bay, South Africa: Perepsilonema benguelae sp. nov. and Leptepsilonema saldanhae sp. nov. (Desmodorida, Epsilonematidae). Zootaxa. 2504: 20–30.
Hendricks, M. G. J.; Gibbons, M. J. (2010). Two new nematode species from Saldanha Bay, South Africa: Perepsilonema benguelae sp. nov. and Leptepsilonema saldanhae sp. nov. (Desmodorida, Epsilonematidae). Zootaxa. 2504: 20–30.