WoRMS source details
Brinkhurst, R.O. (1965). Studies on the North American aquatic Oligochaeta II: Tubificidae. Proceedings of the Academy of Natural Sciences of Philadelphia. 117 (4):117-172.
279532
Brinkhurst, R.O.
1965
Studies on the North American aquatic Oligochaeta II: Tubificidae.
Proceedings of the Academy of Natural Sciences of Philadelphia
117 (4):117-172.
Publication
Available for editors 
[request]
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Forty-four species known from North America are described and figured, and a key is erected for their identification. Most of the species described by earlier American workers (Eisen, Moore, Smith, Verrill) were de-discovered, most of the cosmopolitan species have now been reported, and twelve new species are described. The species may be divided into four geographical groups: - Cosmopolitan, Eastern (with at least related species in Europe), Western (related to Asiatic forms), and Pan- American species. Twenty species are unknown outside North America. Some further new species are now known to occur in the Great Lakes, but these will be described in a separate publication
America, North
Date
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Alexandrovia Hrabĕ, 1962 (additional source)
Aulodrilus Bretscher, 1899 (additional source)
Aulodrilus japonicus Yamaguchi, 1953 (additional source)
Aulodrilus limnobius Bretscher, 1899 (additional source)
Aulodrilus pigueti Kowalewski, 1914 (additional source)
Aulodrilus pluriseta (Piguet, 1906) (additional source)
Bothrioneurum Stolč, 1886 (additional source)
Bothrioneurum vejdovskyanum Štolc, 1886 (additional source)
Branchiura Beddard, 1892 (additional source)
Branchiura sowerbyi Beddard, 1892 (additional source)
Clitellio Savigny, 1822 (additional source)
Clitellio (Clitellio) arenarius (Müller, 1776) (additional source)
Clitellio arenarius (Müller, 1776) represented as Clitellio (Clitellio) arenarius (Müller, 1776) (additional source)
Edmondsonia Brinkhurst, 1965 (original description)
Edmondsonia minor (Sokolskaya, 1961) accepted as Tubifex minor Sokolskaya, 1961 (basis of record)
Edmondsonia montana Brinkhurst, 1965 accepted as Rhyacodrilus montana (Brinkhurst, 1965) (original description)
Euilyodrilus Brinkhurst, 1963 accepted as Potamothrix Vejdovský & Mrázek, 1903 (additional source)
Euilyodrilus bavaricus (Oschmann, 1913) accepted as Potamothrix bavaricus (Oschmann, 1913) (additional source)
Euilyodrilus moldaviensis (Vejdovský & Mrázek, 1903) accepted as Potamothrix moldaviensis Vejdovský & Mrázek, 1903 (additional source)
Ilyodrilus Eisen, 1879 (additional source)
Ilyodrilus asiaticus Chen, 1940 accepted as Chenidrilus asiaticus (Chen, 1940) (additional source)
Ilyodrilus fragilis Eisen, 1879 (additional source)
Ilyodrilus frantzi Brinkhurst, 1965 (original description)
Ilyodrilus frantzi var. capillatus Brinkhurst, 1965 accepted as Ilyodrilus frantzi Brinkhurst, 1965 (original description)
Ilyodrilus frantzi var. typica Brinkhurst, 1965 accepted as Ilyodrilus frantzi Brinkhurst, 1965 (original description)
Ilyodrilus perrieri Eisen, 1879 (additional source)
Isochaeta Pointner, 1911 (additional source)
Isochaeta hamata (Moore, 1905) accepted as Isochaetides hamatus (Moore, 1905) (additional source)
Isochaeta nevadana Brinkhurst, 1965 accepted as Varichaeta nevadana (Brinkhurst, 1965) accepted as Varichaetadrilus minutus (Brinkhurst, 1965) (original description)
Limnodrilus cervix Brinkhurst, 1963 (additional source)
Limnodrilus claparedeanus [subsequent misspelling of 'claparedianus'] accepted as Limnodrilus claparedianus Ratzel, 1868 (basis of record)
Limnodrilus claparedianus Ratzel, 1868 (additional source)
Limnodrilus hoffmeisteri Claparède, 1862 (additional source)
Limnodrilus profundicola (Verrill, 1871) (additional source)
Limnodrilus silvani Eisen, 1879 (additional source)
Limnodrilus udekemianus Claparède, 1862 (additional source)
Monopylephorus Levinsen, 1884 (additional source)
Monopylephorus glaber J. Moore, 1905 accepted as Monopylephorus rubroniveus Levinsen, 1884 (source of synonymy)
Monopylephorus irroratus (Verrill, 1873) (additional source)
Monopylephorus lacteus (Smith, 1900) accepted as Rhizodrilus lacteus Smith, 1900 (basis of record)
Monopylephorus parvus Ditlevsen, 1904 (additional source)
Monopylephorus rubroniveus Levinsen, 1884 (additional source)
Naididae Ehrenberg, 1831 (identification resource)
Peloscolex Leidy, 1852 (additional source)
Peloscolex beetoni Brinkhurst, 1965 accepted as Spirosperma beetoni (Brinkhurst, 1965) (original description)
Peloscolex benedeni (d'Udekem, 1855) accepted as Tubificoides benedii (d'Udekem, 1855) (additional source)
Peloscolex canadensis Brinkhurst, 1965 accepted as Clitellio (Clitellio) arenarius (Müller, 1776) (original description)
Peloscolex carolinensis Brinkhurst, 1965 accepted as Spirosperma carolinensis (Brinkhurst, 1965) (original description)
Peloscolex ferox (Eisen, 1879) accepted as Spirosperma ferox Eisen, 1879 (additional source)
Peloscolex freyi Brinkhurst, 1965 accepted as Isochaetides freyi (Brinkhurst, 1965) (original description)
Peloscolex gabriellae Ev. Marcus, 1950 accepted as Tectidrilus gabriellae (Ev. Marcus, 1950) (additional source)
Peloscolex gabriellae var. apectinata Brinkhurst, 1965 accepted as Tubificoides apectinatus (Brinkhurst, 1965) (original description)
Peloscolex gabriellae var. heterochaetus Brinkhurst, 1965 accepted as Peloscolex gabriellae Ev. Marcus, 1950 accepted as Tectidrilus gabriellae (Ev. Marcus, 1950) (original description)
Peloscolex gabriellae var. nerthoides Brinkhurst, 1965 accepted as Tubificoides nerthoides (Brinkhurst, 1965) (original description)
Peloscolex multisetosus (Smith, 1900) accepted as Quistadrilus multisetosus (Smith, 1900) (additional source)
Peloscolex oregonensis Brinkhurst, 1965 accepted as Embolocephalus nikolskyi (Lastočkin in Lastočkin & Sokolskaja, 1953) (original description)
Peloscolex simsi Brinkhurst, 1966 accepted as Haber speciosus (Hrabĕ, 1931) (taxonomy source)
Peloscolex variegatus Leidy, 1852 (additional source)
Psammoryctides Hrabe, 1964 (additional source)
Psammoryctides californianus Brinkhurst, 1965 (original description)
Psammoryctides minutus Brinkhurst, 1965 accepted as Varichaetadrilus minutus (Brinkhurst, 1965) (original description)
Rhyacodrilus Bretscher, 1901 (additional source)
Rhyacodrilus brevidentatus Brinkhurst, 1965 (original description)
Rhyacodrilus montana (Brinkhurst, 1965) (original description)
Rhyacodrilus sodalis (Eisen, 1879) (additional source)
Spiridion Knöllner, 1935 (additional source)
Spiridion insigne Knöllner, 1935 (additional source)
Spiridion scrobicularae Lastoćkin, 1937 (additional source)
Telmatodrilus Eisen, 1879 (additional source)
Telmatodrilus mcgregori Eisen, 1900 (additional source)
Telmatodrilus vejdovskyi Eisen, 1879 (additional source)
Tubifex Lamarck, 1816 (additional source)
Tubifex longipennis Brinkhurst, 1965 accepted as Tubificoides longipenis (Brinkhurst, 1965) (original description)
Tubifex profundicola Verrill, 1871 accepted as Limnodrilus profundicola (Verrill, 1871) (additional source)
Tubifex pseudogaster (Dahl, 1960) accepted as Tubificoides pseudogaster (Dahl, 1960) (additional source)
Tubifex sinicus Chen, 1940 accepted as Rhyacodrilus sinicus (Chen, 1940) (additional source)
Tubifex templetoni Southern, 1909 accepted as Ilyodrilus templetoni (Southern, 1909) (additional source)
Tubifex tubifex (Müller, 1774) (additional source)
Aulodrilus Bretscher, 1899 (additional source)
Aulodrilus japonicus Yamaguchi, 1953 (additional source)
Aulodrilus limnobius Bretscher, 1899 (additional source)
Aulodrilus pigueti Kowalewski, 1914 (additional source)
Aulodrilus pluriseta (Piguet, 1906) (additional source)
Bothrioneurum Stolč, 1886 (additional source)
Bothrioneurum vejdovskyanum Štolc, 1886 (additional source)
Branchiura Beddard, 1892 (additional source)
Branchiura sowerbyi Beddard, 1892 (additional source)
Clitellio Savigny, 1822 (additional source)
Clitellio (Clitellio) arenarius (Müller, 1776) (additional source)
Clitellio arenarius (Müller, 1776) represented as Clitellio (Clitellio) arenarius (Müller, 1776) (additional source)
Edmondsonia Brinkhurst, 1965 (original description)
Edmondsonia minor (Sokolskaya, 1961) accepted as Tubifex minor Sokolskaya, 1961 (basis of record)
Edmondsonia montana Brinkhurst, 1965 accepted as Rhyacodrilus montana (Brinkhurst, 1965) (original description)
Euilyodrilus Brinkhurst, 1963 accepted as Potamothrix Vejdovský & Mrázek, 1903 (additional source)
Euilyodrilus bavaricus (Oschmann, 1913) accepted as Potamothrix bavaricus (Oschmann, 1913) (additional source)
Euilyodrilus moldaviensis (Vejdovský & Mrázek, 1903) accepted as Potamothrix moldaviensis Vejdovský & Mrázek, 1903 (additional source)
Ilyodrilus Eisen, 1879 (additional source)
Ilyodrilus asiaticus Chen, 1940 accepted as Chenidrilus asiaticus (Chen, 1940) (additional source)
Ilyodrilus fragilis Eisen, 1879 (additional source)
Ilyodrilus frantzi Brinkhurst, 1965 (original description)
Ilyodrilus frantzi var. capillatus Brinkhurst, 1965 accepted as Ilyodrilus frantzi Brinkhurst, 1965 (original description)
Ilyodrilus frantzi var. typica Brinkhurst, 1965 accepted as Ilyodrilus frantzi Brinkhurst, 1965 (original description)
Ilyodrilus perrieri Eisen, 1879 (additional source)
Isochaeta Pointner, 1911 (additional source)
Isochaeta hamata (Moore, 1905) accepted as Isochaetides hamatus (Moore, 1905) (additional source)
Isochaeta nevadana Brinkhurst, 1965 accepted as Varichaeta nevadana (Brinkhurst, 1965) accepted as Varichaetadrilus minutus (Brinkhurst, 1965) (original description)
Limnodrilus cervix Brinkhurst, 1963 (additional source)
Limnodrilus claparedeanus [subsequent misspelling of 'claparedianus'] accepted as Limnodrilus claparedianus Ratzel, 1868 (basis of record)
Limnodrilus claparedianus Ratzel, 1868 (additional source)
Limnodrilus hoffmeisteri Claparède, 1862 (additional source)
Limnodrilus profundicola (Verrill, 1871) (additional source)
Limnodrilus silvani Eisen, 1879 (additional source)
Limnodrilus udekemianus Claparède, 1862 (additional source)
Monopylephorus Levinsen, 1884 (additional source)
Monopylephorus glaber J. Moore, 1905 accepted as Monopylephorus rubroniveus Levinsen, 1884 (source of synonymy)
Monopylephorus irroratus (Verrill, 1873) (additional source)
Monopylephorus lacteus (Smith, 1900) accepted as Rhizodrilus lacteus Smith, 1900 (basis of record)
Monopylephorus parvus Ditlevsen, 1904 (additional source)
Monopylephorus rubroniveus Levinsen, 1884 (additional source)
Naididae Ehrenberg, 1831 (identification resource)
Peloscolex Leidy, 1852 (additional source)
Peloscolex beetoni Brinkhurst, 1965 accepted as Spirosperma beetoni (Brinkhurst, 1965) (original description)
Peloscolex benedeni (d'Udekem, 1855) accepted as Tubificoides benedii (d'Udekem, 1855) (additional source)
Peloscolex canadensis Brinkhurst, 1965 accepted as Clitellio (Clitellio) arenarius (Müller, 1776) (original description)
Peloscolex carolinensis Brinkhurst, 1965 accepted as Spirosperma carolinensis (Brinkhurst, 1965) (original description)
Peloscolex ferox (Eisen, 1879) accepted as Spirosperma ferox Eisen, 1879 (additional source)
Peloscolex freyi Brinkhurst, 1965 accepted as Isochaetides freyi (Brinkhurst, 1965) (original description)
Peloscolex gabriellae Ev. Marcus, 1950 accepted as Tectidrilus gabriellae (Ev. Marcus, 1950) (additional source)
Peloscolex gabriellae var. apectinata Brinkhurst, 1965 accepted as Tubificoides apectinatus (Brinkhurst, 1965) (original description)
Peloscolex gabriellae var. heterochaetus Brinkhurst, 1965 accepted as Peloscolex gabriellae Ev. Marcus, 1950 accepted as Tectidrilus gabriellae (Ev. Marcus, 1950) (original description)
Peloscolex gabriellae var. nerthoides Brinkhurst, 1965 accepted as Tubificoides nerthoides (Brinkhurst, 1965) (original description)
Peloscolex multisetosus (Smith, 1900) accepted as Quistadrilus multisetosus (Smith, 1900) (additional source)
Peloscolex oregonensis Brinkhurst, 1965 accepted as Embolocephalus nikolskyi (Lastočkin in Lastočkin & Sokolskaja, 1953) (original description)
Peloscolex simsi Brinkhurst, 1966 accepted as Haber speciosus (Hrabĕ, 1931) (taxonomy source)
Peloscolex variegatus Leidy, 1852 (additional source)
Psammoryctides Hrabe, 1964 (additional source)
Psammoryctides californianus Brinkhurst, 1965 (original description)
Psammoryctides minutus Brinkhurst, 1965 accepted as Varichaetadrilus minutus (Brinkhurst, 1965) (original description)
Rhyacodrilus Bretscher, 1901 (additional source)
Rhyacodrilus brevidentatus Brinkhurst, 1965 (original description)
Rhyacodrilus montana (Brinkhurst, 1965) (original description)
Rhyacodrilus sodalis (Eisen, 1879) (additional source)
Spiridion Knöllner, 1935 (additional source)
Spiridion insigne Knöllner, 1935 (additional source)
Spiridion scrobicularae Lastoćkin, 1937 (additional source)
Telmatodrilus Eisen, 1879 (additional source)
Telmatodrilus mcgregori Eisen, 1900 (additional source)
Telmatodrilus vejdovskyi Eisen, 1879 (additional source)
Tubifex Lamarck, 1816 (additional source)
Tubifex longipennis Brinkhurst, 1965 accepted as Tubificoides longipenis (Brinkhurst, 1965) (original description)
Tubifex profundicola Verrill, 1871 accepted as Limnodrilus profundicola (Verrill, 1871) (additional source)
Tubifex pseudogaster (Dahl, 1960) accepted as Tubificoides pseudogaster (Dahl, 1960) (additional source)
Tubifex sinicus Chen, 1940 accepted as Rhyacodrilus sinicus (Chen, 1940) (additional source)
Tubifex templetoni Southern, 1909 accepted as Ilyodrilus templetoni (Southern, 1909) (additional source)
Tubifex tubifex (Müller, 1774) (additional source)
Holotype USNM 25670, verbatimGeounit Flag Lake, Havana, I..., identified as Peloscolex multisetosus (Smith, 1900)
Holotype USNM 32599, verbatimGeounit Under stones, Long B..., identified as Peloscolex canadensis Brinkhurst, 1965
Holotype USNM 32606, verbatimGeounit Five Islands, Dry Po..., identified as Tubifex longipenis Brinkhurst, 1965
Holotype USNM 32620, identified as Peloscolex beetoni Brinkhurst, 1965
Holotype USNM 32623, verbatimGeounit Berry Creek, Corvali..., identified as Peloscolex oregonensis Brinkhurst, 1965
Neotype USNM 32626, identified as Peloscolex variegatus Leidy, 1852
Holotype USNM 32628, verbatimGeounit White Lake, North Ca..., identified as Peloscolex freyi Brinkhurst, 1965
Holotype USNM 32632, verbatimGeounit White Lake, North Ca..., identified as Peloscolex carolinensis Brinkhurst, 1965
Holotype USNM 32639, identified as Psammoryctides minutus Brinkhurst, 1965
Holotype USNM 32642, identified as Isochaeta nevadana Brinkhurst, 1965
Holotype USNM 32646, identified as Rhyacodrilus brevidentatus Brinkhurst, 1965
Holotype USNM 32652, verbatimGeounit Dog Lake, Yakima Cou..., identified as Edmondsonia montana Brinkhurst, 1965
Holotype USNM 32655, identified as Ilyodrilus frantzi Brinkhurst, 1965
Holotype USNM 32662, verbatimGeounit Coyote Creek, Santa ..., identified as Psammoryctides californianus Brinkhurst, 1965
Holotype USNM 32599, verbatimGeounit Under stones, Long B..., identified as Peloscolex canadensis Brinkhurst, 1965
Holotype USNM 32606, verbatimGeounit Five Islands, Dry Po..., identified as Tubifex longipenis Brinkhurst, 1965
Holotype USNM 32620, identified as Peloscolex beetoni Brinkhurst, 1965
Holotype USNM 32623, verbatimGeounit Berry Creek, Corvali..., identified as Peloscolex oregonensis Brinkhurst, 1965
Neotype USNM 32626, identified as Peloscolex variegatus Leidy, 1852
Holotype USNM 32628, verbatimGeounit White Lake, North Ca..., identified as Peloscolex freyi Brinkhurst, 1965
Holotype USNM 32632, verbatimGeounit White Lake, North Ca..., identified as Peloscolex carolinensis Brinkhurst, 1965
Holotype USNM 32639, identified as Psammoryctides minutus Brinkhurst, 1965
Holotype USNM 32642, identified as Isochaeta nevadana Brinkhurst, 1965
Holotype USNM 32646, identified as Rhyacodrilus brevidentatus Brinkhurst, 1965
Holotype USNM 32652, verbatimGeounit Dog Lake, Yakima Cou..., identified as Edmondsonia montana Brinkhurst, 1965
Holotype USNM 32655, identified as Ilyodrilus frantzi Brinkhurst, 1965
Holotype USNM 32662, verbatimGeounit Coyote Creek, Santa ..., identified as Psammoryctides californianus Brinkhurst, 1965
Etymology
The species is named for my (edit. R.O. Brinkhurst) colleague, Dr. A. M. Beeton. [details]
Nomenclature
Psammoryctes Vejdovsky, 1875 — name amended Hrabë (1964): Publ. Fac. Univ. Brno 450: 101-112. [details]
Spelling
Limnodrilus claparedianus Ratzel is named in honor of Swiss Professor René-Edouard Claparède (1832-1871). The ... [details]
