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Rouse, Greg W.; Carvajal, Jose Ignacio; Pleijel, Fredrik. (2018). Phylogeny of Hesionidae (Aciculata, Annelida), with four new species from deep-sea eastern Pacific methane seeps, and resolution of the affinity of Hesiolyra. Invertebrate Systematics. 32(5): 1050-1068.
Rouse, Greg W.; Carvajal, Jose Ignacio; Pleijel, Fredrik. (2018). Phylogeny of Hesionidae (Aciculata, Annelida), with four new species from deep-sea eastern Pacific methane seeps, and resolution of the affinity of Hesiolyra. Invertebrate Systematics. 32(5): 1050-1068.