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Apel, M.; Spiridonov, V. A. (1998). Taxonomy and zoogeography of the portunid crabs (Crustacea: Decapoda: Brachyura: Portunidae) of the Arabian Gulf and adjacent waters. Fauna of Arabia. 17: 159-331.
Apel, M.; Spiridonov, V. A. (1998). Taxonomy and zoogeography of the portunid crabs (Crustacea: Decapoda: Brachyura: Portunidae) of the Arabian Gulf and adjacent waters. Fauna of Arabia. 17: 159-331.