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Hsu, J.-W.; Shih, H.-T. (2020). Diversity of Taiwanese Brackish Crabs Genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) based on DNA Barcodes, with Descriptions of Two New Species. Zoological Studies. 59: 59.
Hsu, J.-W.; Shih, H.-T. (2020). Diversity of Taiwanese Brackish Crabs Genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) based on DNA Barcodes, with Descriptions of Two New Species. Zoological Studies. 59: 59.
